Links
-Google scholar page here
-Open access versions of most papers are available here
-Preprints can be found here
-Open Science Framework page here
-Orcid page: here
2024
60. Aves, Kornysheva, Galea. Holistic planning of sequential movements. In prep
59. Sporn, Galea. Reward-based improvements in motor sequence learning are differentially affected by dopamine. Under review (bioRxiv)
58. Dawson, Galea, Sporn, Krakauer, Bestmann, Ward. High-dose high-intensity neurorehabilitation in chronic stroke improves general arm motor control. Under review
57. Hickman, Sowden, Fraser, Schuster, Rybicki, Galea, Cook. Dopaminergic manipulations affect the modulation and meta-modulation of movement speed: evidence from two pharmacological interventions. Behavioural Brain Research.
56. Hamel, Walzing, Hinder, McAllister, Jenkinson, Galea. Bilateral intracortical inhibition during unilateral motor preparation and sequence learning. Brain Stimulation
2023
55. Hamel, Pearson, Sifi, Patel, Hinder, Jenkinson, Galea. The intracortical excitability changes underlying the enhancing effects of rewards and punishment on motor performance. Brain Stimulation
54. Weightman, Lalji, Lin, Galea, Jenkinson & Miall. Short duration event-related tdcs enhances visuomotor adaptation. Brain Stimulation
53. Codol, Kashefi, Forgaard, Galea, Pruszyski & Gribble. Sensorimotor feedback loops are selectively sensitive to reward. eLife
52. Abdlkarim, Di Luca, Aves, Yeo, Miall, Holland & Galea. A methodological framework to assess the accuracy of virtual reality hand-tracking systems. Behavior Research Methods
2022
51. Sporn, Chen & Galea. The dissociable effects of reward on sequential motor behaviour. Journal of Neurophysiology
2021
50. Rybicki, Galea, Hiles, Fabin & Cook. Intact predictive motor sequence learning in autism spectrum disorder. Scientific Reports
2020
49. Codol, Galea, Jalali & Holland. Reward-driven enhancements in motor control are robust to TMS manipulation. Experimental Brain Research
48. Codol, Holland, Manohar & Galea. Reward-based improvements in motor control are driven by multiple error-reducing mechanisms. Journal of Neuroscience
47. Chen, Voet, Jenkinson & Galea. Dopamine-dependent loss aversion during effort-based decision-making. Journal of Neuroscience
2019
46. Holland*, Codol*, Oxley, Taylor, Hamshere, Joseph & Galea. Domain-specific working memory, but not dopamine-related genetic variability, shapes reward-based motor learning. Journal of Neuroscience *Equal contribution
45. Sadnicka, Galea & Edwards. What can we kinematic studies tell us about the mechanism of dystonia? Progress in Brain Research
2018
44. Sadnicka, Galea, Chen, Warner, Bhatia, Rothwell & Edwards. Delineating cerebellar mechanisms in DYT11 myoclonus-dystonia. Movement Disorders
43. Chen X, Rutledge R, Brown H, Dolan R, Bestmann S* & Galea*. Age-dependent Pavlovian biases influence motor decision-making. PLOS Computational Biology * equal contribution.
42. Codol O, Holland P & Galea. The relationship between reinforcement and explicit control during visuomotor adaptation. Nature Scientific Reports
41. Quattrocchi G, Monaco, Ho, Irmen, Strube, Ruge D, Bestmann S* & Galea*. Pharmacological dopamine manipulation does not alter reward-based improvements in memory retention during a visuomotor adaptation task. eNeuro * equal contribution.
40. Holland P, Codol O & Galea. The contribution of explicit processes to reinforcement-based motor learning. Journal of Neurophysiology
39. Sadnicka A, Stevenson A, Bhatia K, Rothwell JC, Edwards M & Galea. High motor variability in DYT1 dystonia is associated with impaired visuomotor adaptation. Nature Scientific Reports
38. Jalali R, Chowdhury A, Wilson M, Miall RC & Galea. Neural changes associated with cerebellar tDCS studied using MR spectroscopy. Experimental Brain Research.
2017
37. Chen X*, Holland P* & Galea. The effects of reward and punishment on motor skill learning. Current Opinion in Behavioral Science. * equal contribution. Click for manuscript
36. Chen X, Mohr K & Galea. Predicting explorative motor learning using decision-making and motor noise. PLOS Computational Biology.
35. Jalali R, Miall RC & Galea. No consistent effect of cerebellar tDCS on visuomotor adaptation. Journal of Neurophysiology. Click for manuscript
34. Quattrocchi G, Greenwood R, Rothwell JC, Galea*, Bestmann S*. Reward and Punishment enhance motor adaptation in stroke. JNNP * equal contribution.
2016
33. Christou AI, Miall RC, McNab F* & Galea*. Individual differences in explicit and implicit visuomotor learning and working memory capacity. Nature Scientific Reports 6:36633. * equal contribution.
32. Mevorach C, Spaniol MM, Soden M & Galea. Age-dependent distractor suppression across the vision and motor domain. Journal of Vision 16(11):27
31. Caligiore D, Pezzulo G, Baldassarre G, Bostan AC, Strick PL, Doya K, Helmich RC, Dirkx M, Houk J, Jörntell H, Lago-Rodriguez A, Galea, Miall RC, Popa T, Kishore A, Verschure P, Zucca R, Herreros I. Consensus Paper: Towards a Systems-Level View of Cerebellar Function: the Interplay Between Cerebellum, Basal Ganglia, and Cortex. Cerebellum 16(1):203-229.
30. Miall RC, Galea. Cerebellar damage limits reinforcement learning. Commentary on Therrien et al: Effective reinforcement learning following cerebellar damage requires a balance between exploration and motor noise. Brain 139(1):4-7.
2015
29. Tomassini A, Ruge D, Galea, Penny W, Bestmann S. The role of dopamine in temporal uncertainty. Journal of Cognitive Neuroscience 28(1):597-602.
28. Galea, Mallia E, Rothwell JC, Diedrichsen J. The dissociable effects of punishment and reward on motor learning. Nature Neuroscience 18(4): 597-602.
27. Schlerf, J, Galea, Spanpinato, D, Celnik, P (2014). Laterality differences in cerebellar connectivity revealed by TMS. Cerebral Cortex 25(7):1827-34.
2014
26. Grimaldi G, Argyropoulos G, Bastian A, Cortes M, Davis N, Edwards D, Ferrucci D, Fregni F, Galea, Hamada M, Manto M, Miall R, Morales-Quezada L, Pope P, Priori A, Rothwell J, Tomlinson S, Celnik P. Cerebellar Transcranial Direct Current Stimulation (ctDCS): A Novel Approach To Understanding Cerebellar Function In Health And Disease. Neuroscientist 22(1):83-97.
25. Hamada M, Galea, Lazzaro V, Mazzone M, Ziemann U, Rothwell JC. The role of two separate motor cortex interneuron circuits in different forms of human physiological and behavioural plasticity. Journal of Neuroscience 34(38):12837-49.
24. Bestmann S, Ruge R, Rothwell JC, Galea. The role of dopamine in motor flexibility. Journal of Cognitive Neuroscience 27(2):365-76.
23. Sadnicka A, Patani B, Saifee T, Kassavetis P, Pareés I, Korlipara P, Bhatia KP, Rothwell JC, Galea*, Edwards MJ*. Normal motor adaptation in cervical dystonia: a fundamental cerebellar computation is intact. Cerebellum 13(5):535-44. * equal contribution.
22. Salas RE, Galea, Spira AP, Bremers A, Walston J, Celnik P. Increased use-dependent plasticity in chronic insomnia. Sleep 37(3):535-44.
2013
21. Grimaldi G, Argyropoulos GP, Boehringer A, Celnik P, Edwards MJ, Ferrucci R, Galea, Groiss SJ, Hiraoka K, Kassavetis P, Lesage L, Manto M, Miall RC, Priori A, Sadnicka A, Ugawa Y, Ziemann U. Cerebellar neurostimulation - a consensus paper. Cerebellum 13(1):121-38.
20. Sadnicka A, Kimmich O, Pisarek C, Ruge R, Galea, Kassavetis P, Pareés I, Saifee T, Molloy A, Bradley D, O'Riordan S, Zrinzo L, Hariz M, Bhatia K, Limousin P, Foltynie T, Rothwell J, Hutchinson M, Edwards M. Pallidal stimulation for cervical dystonia does not correct abnormal temporal discrimination. Movement Disorders 13(1):121-38..
19. Galea, Ruge D, Buijink A, Bestmann S & Rothwell J.C. Punishment induced behavioural and neurophysiological variability reveals dopamine-dependent selection of kinematic movement parameters. Journal of Neuroscience 33(9):3981-8.
18. Hasan A, Galea, Casula E.P, Falkai P, Bestmann S & Rothwell J.C. Muscle and timing-specific functional connectivity between the dorsolateral prefrontal cortex and the primary motor cortex. Journal of Cognitive Neuroscience 25(4):558-70.
2012
17. Schlerf J, Galea, Bastian A.J & Celnik P. Dynamic modulation of cerebellar excitability for abrupt, but not gradual visuomotor adaptation. Journal of Neuroscience 32(34):11610-7.
16. Hamada M, Strigaro G, Murase N, Sadnicka A, Galea, Edwards M.J & Rothwell J.C. Cerebellar modulation of human associative plasticity. Journal of Physiology 590(10):2365-74.
15. Friston K, Shiner T, Fitzger T, Galea, Adams R, Brown H, Dolan R, Moran R, Stephan K & Bestmann S. Dopamine, precision and affordance in active inference. PLOS Computational Biology 8(1).
14. Hasan A, Wobrock T, Falkai P, Schneider-Axmann T, Guse B, Backens M, Ecker U.K.H, Heimes J, Galea, Gruber O, Scherk H. Hippocampal integrity and neurocognition in first–episode schizophrenia: a multidimensional study. The World Journal of Biological Psychiatry 15(3):188-99.
13. Hasan A, Hamada M, Nitsche N.A, Ruge D, Galea, Wobrock T & Rothwell J.C. Direct-current dependent shift of theta-burst induced plasticity in the human motor cortex. Experimental Brain Research 217(1):15-23.
12. Galea*, Bestmann S*, Beigi M, Jahanshahi M & Rothwell J.C. Action reprogramming in Parkinson’s disease: response to prediction errors is modulated by levels of dopamine. Journal of Neuroscience 32(2):542-50. * equal contribution.
2011
11. Cantarero G, Galea, Ajagbe L, Salas R, Willis J & Celnik P. Disrupting the Ventral Premotor Cortex Interferes with the Contribution of Action Observation to Use-dependent Plasticity. Journal of Cognitive Neuroscience 23(12):3757-66.
10. Holland R, Leff A.P, Joseph O, Galea, Mahalekshmi D, Price C.J, Rothwell J.C & Crinion J. Left frontal anodal tDCS elicits reduced neural activity in Broca’s area and faster naming responses. Current Biology 21(16):1403-7.
9. Jayaram G, Galea, Bastian A.J & Celnik P. Human locomotor adaptive learning is proportional to depression of cerebellar excitability. Cerebral Cortex 21(8):1901-9.
8. Galea, Vazquez A, Pasricha N, Orban de Xivry J.J & Celnik P. Dissociating the roles of the cerebellum and motor cortex during adaptive learning: the motor cortex retains what the cerebellum learns. Cerebral Cortex 21(8):1761-70.
2010
7. Galea, Saber S, Albert N.B & Miall R.C. Secondary tasks impair adaptation to step and gradual visual displacements. Experimental Brain Research 202(2):473:84.
6. Galea*, Albert N.B*, Dyte T & Miall R.C. Theta-burst TMS to the DLPFC facilitates the consolidation of procedural skills. Journal of Cognitive Neuroscience 22(6):1158-68. * equal contribution.
2009
5. Galea, Jayaram G, Ajagbe L & Celnik P. Modulation of cerebellar excitability by polarity-specific non-invasive direct current stimulation. Journal of Neuroscience 29:9115-22.
4. Galea & Celnik P. Brain polarization enhances the formation and retention of motor memories. Journal of Neurophysiology 102: 294-301.
2007
3. Galea, Woolley D & Miall R.C. Asymmetric interlimb transfer of concurrent adaptation to opposing dynamic forces. Experimental Brain Research 182:267-273.
2. Jackson C, Albert N, Roberts R, Galea & Swait G. Target selection: choice or response? Journal of Neuroscience 27:6079-6080.
2006
1. Galea & Miall R.C. Concurrent adaptation to opposing visual displacements during an alternating movement. Experimental Brain Research 175(4):676-688.
* = Authors contributed equally to this work.
-Google scholar page here
-Open access versions of most papers are available here
-Preprints can be found here
-Open Science Framework page here
-Orcid page: here
2024
60. Aves, Kornysheva, Galea. Holistic planning of sequential movements. In prep
59. Sporn, Galea. Reward-based improvements in motor sequence learning are differentially affected by dopamine. Under review (bioRxiv)
58. Dawson, Galea, Sporn, Krakauer, Bestmann, Ward. High-dose high-intensity neurorehabilitation in chronic stroke improves general arm motor control. Under review
57. Hickman, Sowden, Fraser, Schuster, Rybicki, Galea, Cook. Dopaminergic manipulations affect the modulation and meta-modulation of movement speed: evidence from two pharmacological interventions. Behavioural Brain Research.
56. Hamel, Walzing, Hinder, McAllister, Jenkinson, Galea. Bilateral intracortical inhibition during unilateral motor preparation and sequence learning. Brain Stimulation
2023
55. Hamel, Pearson, Sifi, Patel, Hinder, Jenkinson, Galea. The intracortical excitability changes underlying the enhancing effects of rewards and punishment on motor performance. Brain Stimulation
54. Weightman, Lalji, Lin, Galea, Jenkinson & Miall. Short duration event-related tdcs enhances visuomotor adaptation. Brain Stimulation
53. Codol, Kashefi, Forgaard, Galea, Pruszyski & Gribble. Sensorimotor feedback loops are selectively sensitive to reward. eLife
52. Abdlkarim, Di Luca, Aves, Yeo, Miall, Holland & Galea. A methodological framework to assess the accuracy of virtual reality hand-tracking systems. Behavior Research Methods
2022
51. Sporn, Chen & Galea. The dissociable effects of reward on sequential motor behaviour. Journal of Neurophysiology
2021
50. Rybicki, Galea, Hiles, Fabin & Cook. Intact predictive motor sequence learning in autism spectrum disorder. Scientific Reports
2020
49. Codol, Galea, Jalali & Holland. Reward-driven enhancements in motor control are robust to TMS manipulation. Experimental Brain Research
48. Codol, Holland, Manohar & Galea. Reward-based improvements in motor control are driven by multiple error-reducing mechanisms. Journal of Neuroscience
47. Chen, Voet, Jenkinson & Galea. Dopamine-dependent loss aversion during effort-based decision-making. Journal of Neuroscience
2019
46. Holland*, Codol*, Oxley, Taylor, Hamshere, Joseph & Galea. Domain-specific working memory, but not dopamine-related genetic variability, shapes reward-based motor learning. Journal of Neuroscience *Equal contribution
45. Sadnicka, Galea & Edwards. What can we kinematic studies tell us about the mechanism of dystonia? Progress in Brain Research
2018
44. Sadnicka, Galea, Chen, Warner, Bhatia, Rothwell & Edwards. Delineating cerebellar mechanisms in DYT11 myoclonus-dystonia. Movement Disorders
43. Chen X, Rutledge R, Brown H, Dolan R, Bestmann S* & Galea*. Age-dependent Pavlovian biases influence motor decision-making. PLOS Computational Biology * equal contribution.
42. Codol O, Holland P & Galea. The relationship between reinforcement and explicit control during visuomotor adaptation. Nature Scientific Reports
41. Quattrocchi G, Monaco, Ho, Irmen, Strube, Ruge D, Bestmann S* & Galea*. Pharmacological dopamine manipulation does not alter reward-based improvements in memory retention during a visuomotor adaptation task. eNeuro * equal contribution.
40. Holland P, Codol O & Galea. The contribution of explicit processes to reinforcement-based motor learning. Journal of Neurophysiology
39. Sadnicka A, Stevenson A, Bhatia K, Rothwell JC, Edwards M & Galea. High motor variability in DYT1 dystonia is associated with impaired visuomotor adaptation. Nature Scientific Reports
38. Jalali R, Chowdhury A, Wilson M, Miall RC & Galea. Neural changes associated with cerebellar tDCS studied using MR spectroscopy. Experimental Brain Research.
2017
37. Chen X*, Holland P* & Galea. The effects of reward and punishment on motor skill learning. Current Opinion in Behavioral Science. * equal contribution. Click for manuscript
36. Chen X, Mohr K & Galea. Predicting explorative motor learning using decision-making and motor noise. PLOS Computational Biology.
35. Jalali R, Miall RC & Galea. No consistent effect of cerebellar tDCS on visuomotor adaptation. Journal of Neurophysiology. Click for manuscript
34. Quattrocchi G, Greenwood R, Rothwell JC, Galea*, Bestmann S*. Reward and Punishment enhance motor adaptation in stroke. JNNP * equal contribution.
2016
33. Christou AI, Miall RC, McNab F* & Galea*. Individual differences in explicit and implicit visuomotor learning and working memory capacity. Nature Scientific Reports 6:36633. * equal contribution.
32. Mevorach C, Spaniol MM, Soden M & Galea. Age-dependent distractor suppression across the vision and motor domain. Journal of Vision 16(11):27
31. Caligiore D, Pezzulo G, Baldassarre G, Bostan AC, Strick PL, Doya K, Helmich RC, Dirkx M, Houk J, Jörntell H, Lago-Rodriguez A, Galea, Miall RC, Popa T, Kishore A, Verschure P, Zucca R, Herreros I. Consensus Paper: Towards a Systems-Level View of Cerebellar Function: the Interplay Between Cerebellum, Basal Ganglia, and Cortex. Cerebellum 16(1):203-229.
30. Miall RC, Galea. Cerebellar damage limits reinforcement learning. Commentary on Therrien et al: Effective reinforcement learning following cerebellar damage requires a balance between exploration and motor noise. Brain 139(1):4-7.
2015
29. Tomassini A, Ruge D, Galea, Penny W, Bestmann S. The role of dopamine in temporal uncertainty. Journal of Cognitive Neuroscience 28(1):597-602.
28. Galea, Mallia E, Rothwell JC, Diedrichsen J. The dissociable effects of punishment and reward on motor learning. Nature Neuroscience 18(4): 597-602.
27. Schlerf, J, Galea, Spanpinato, D, Celnik, P (2014). Laterality differences in cerebellar connectivity revealed by TMS. Cerebral Cortex 25(7):1827-34.
2014
26. Grimaldi G, Argyropoulos G, Bastian A, Cortes M, Davis N, Edwards D, Ferrucci D, Fregni F, Galea, Hamada M, Manto M, Miall R, Morales-Quezada L, Pope P, Priori A, Rothwell J, Tomlinson S, Celnik P. Cerebellar Transcranial Direct Current Stimulation (ctDCS): A Novel Approach To Understanding Cerebellar Function In Health And Disease. Neuroscientist 22(1):83-97.
25. Hamada M, Galea, Lazzaro V, Mazzone M, Ziemann U, Rothwell JC. The role of two separate motor cortex interneuron circuits in different forms of human physiological and behavioural plasticity. Journal of Neuroscience 34(38):12837-49.
24. Bestmann S, Ruge R, Rothwell JC, Galea. The role of dopamine in motor flexibility. Journal of Cognitive Neuroscience 27(2):365-76.
23. Sadnicka A, Patani B, Saifee T, Kassavetis P, Pareés I, Korlipara P, Bhatia KP, Rothwell JC, Galea*, Edwards MJ*. Normal motor adaptation in cervical dystonia: a fundamental cerebellar computation is intact. Cerebellum 13(5):535-44. * equal contribution.
22. Salas RE, Galea, Spira AP, Bremers A, Walston J, Celnik P. Increased use-dependent plasticity in chronic insomnia. Sleep 37(3):535-44.
2013
21. Grimaldi G, Argyropoulos GP, Boehringer A, Celnik P, Edwards MJ, Ferrucci R, Galea, Groiss SJ, Hiraoka K, Kassavetis P, Lesage L, Manto M, Miall RC, Priori A, Sadnicka A, Ugawa Y, Ziemann U. Cerebellar neurostimulation - a consensus paper. Cerebellum 13(1):121-38.
20. Sadnicka A, Kimmich O, Pisarek C, Ruge R, Galea, Kassavetis P, Pareés I, Saifee T, Molloy A, Bradley D, O'Riordan S, Zrinzo L, Hariz M, Bhatia K, Limousin P, Foltynie T, Rothwell J, Hutchinson M, Edwards M. Pallidal stimulation for cervical dystonia does not correct abnormal temporal discrimination. Movement Disorders 13(1):121-38..
19. Galea, Ruge D, Buijink A, Bestmann S & Rothwell J.C. Punishment induced behavioural and neurophysiological variability reveals dopamine-dependent selection of kinematic movement parameters. Journal of Neuroscience 33(9):3981-8.
18. Hasan A, Galea, Casula E.P, Falkai P, Bestmann S & Rothwell J.C. Muscle and timing-specific functional connectivity between the dorsolateral prefrontal cortex and the primary motor cortex. Journal of Cognitive Neuroscience 25(4):558-70.
2012
17. Schlerf J, Galea, Bastian A.J & Celnik P. Dynamic modulation of cerebellar excitability for abrupt, but not gradual visuomotor adaptation. Journal of Neuroscience 32(34):11610-7.
16. Hamada M, Strigaro G, Murase N, Sadnicka A, Galea, Edwards M.J & Rothwell J.C. Cerebellar modulation of human associative plasticity. Journal of Physiology 590(10):2365-74.
15. Friston K, Shiner T, Fitzger T, Galea, Adams R, Brown H, Dolan R, Moran R, Stephan K & Bestmann S. Dopamine, precision and affordance in active inference. PLOS Computational Biology 8(1).
14. Hasan A, Wobrock T, Falkai P, Schneider-Axmann T, Guse B, Backens M, Ecker U.K.H, Heimes J, Galea, Gruber O, Scherk H. Hippocampal integrity and neurocognition in first–episode schizophrenia: a multidimensional study. The World Journal of Biological Psychiatry 15(3):188-99.
13. Hasan A, Hamada M, Nitsche N.A, Ruge D, Galea, Wobrock T & Rothwell J.C. Direct-current dependent shift of theta-burst induced plasticity in the human motor cortex. Experimental Brain Research 217(1):15-23.
12. Galea*, Bestmann S*, Beigi M, Jahanshahi M & Rothwell J.C. Action reprogramming in Parkinson’s disease: response to prediction errors is modulated by levels of dopamine. Journal of Neuroscience 32(2):542-50. * equal contribution.
2011
11. Cantarero G, Galea, Ajagbe L, Salas R, Willis J & Celnik P. Disrupting the Ventral Premotor Cortex Interferes with the Contribution of Action Observation to Use-dependent Plasticity. Journal of Cognitive Neuroscience 23(12):3757-66.
10. Holland R, Leff A.P, Joseph O, Galea, Mahalekshmi D, Price C.J, Rothwell J.C & Crinion J. Left frontal anodal tDCS elicits reduced neural activity in Broca’s area and faster naming responses. Current Biology 21(16):1403-7.
9. Jayaram G, Galea, Bastian A.J & Celnik P. Human locomotor adaptive learning is proportional to depression of cerebellar excitability. Cerebral Cortex 21(8):1901-9.
8. Galea, Vazquez A, Pasricha N, Orban de Xivry J.J & Celnik P. Dissociating the roles of the cerebellum and motor cortex during adaptive learning: the motor cortex retains what the cerebellum learns. Cerebral Cortex 21(8):1761-70.
2010
7. Galea, Saber S, Albert N.B & Miall R.C. Secondary tasks impair adaptation to step and gradual visual displacements. Experimental Brain Research 202(2):473:84.
6. Galea*, Albert N.B*, Dyte T & Miall R.C. Theta-burst TMS to the DLPFC facilitates the consolidation of procedural skills. Journal of Cognitive Neuroscience 22(6):1158-68. * equal contribution.
2009
5. Galea, Jayaram G, Ajagbe L & Celnik P. Modulation of cerebellar excitability by polarity-specific non-invasive direct current stimulation. Journal of Neuroscience 29:9115-22.
4. Galea & Celnik P. Brain polarization enhances the formation and retention of motor memories. Journal of Neurophysiology 102: 294-301.
2007
3. Galea, Woolley D & Miall R.C. Asymmetric interlimb transfer of concurrent adaptation to opposing dynamic forces. Experimental Brain Research 182:267-273.
2. Jackson C, Albert N, Roberts R, Galea & Swait G. Target selection: choice or response? Journal of Neuroscience 27:6079-6080.
2006
1. Galea & Miall R.C. Concurrent adaptation to opposing visual displacements during an alternating movement. Experimental Brain Research 175(4):676-688.
* = Authors contributed equally to this work.